When the Museum of Natural History is closed for upgrades and renovations, the museum pieces are moved into federal storage at the Smithsonian Institution, which comes to life. The museum houses the world's largest museum complex with more than 136 million items in its collections, ranging from the plane Amelia Earhart (Amy Adams) flew on her non-stop solo flight across the Atlantic and Al Capone's (Jon Bernthal) rap sheet and mug shot to Dorothy's ruby slippers, Fonzie's jacket from Happy Days, the still from M*A*S*H and Archie Bunker's lounge chair from All in the Family. When the museum pieces from the first film are attacked by Kahmunrah, an evil Pharaoh, Larry (Ben Stiller) breaks into the Smithsonian to save the old exhibits from New York (Such as Theodore Roosevelt and Dexter) from new exhibits (like General Custer, Napoleon Bonaparte, Al Capone, and even Darth Vader) (and this means they supported Kahmunrah). Larry enlists the help of Amelia Earhart, whom he develops a romantic interest in, and together they try to put everything back in order.
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